Cerioidini

Cerioidini
	Ceriana vespiformes
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Arthropoda
Class:	Insecta
Order:	Diptera
Family:	Syrphidae
Subfamily:	Eristalinae
Tribe:	Cerioidini; Wahlgren, 1909
Genera
	Ceriana Rafinesque, 1815; Monoceromyia Shannon, 1925; Polybiomyia Shannon, 1925; Primocerioides; Sphiximorpha Rondani 1850;

Cerioidini is a widespread tribe of around 222 species of hoverfly.^[1] Cerioidini are mistaken for wasps for which they are effective mimic. Cerioidini have antennae with a terminal style and have somewhat elongate and basally constricted abdomens, only slightly in Ceriana, but pronounced in most Sphiximorpha; and Polybiomyia, and extremely in Monoceromyia. ^[2] Larvae live mostly within tree sap associated with tree wounds or putrefying pockets of water in tree cavities. ^[3]

Description

This family is distinguished by anatomy and coloration that has evolved to be wasp mimics. The antennae are long and thin with a terminal arista (see gallery) suggestive of wasp antennae that are also long. The arista of the antennae is terminal. The elongate antennae has arisen independently in other genera such as Mocrodon although in that genera the arista is not terminal. The flagellum is always elongate while the scape and pedicel are elongated in different ratios that are helpful in distinguishing the various genera. an elongation of the front, the frontal prominence adds to the elongation of the antennae but is reduced or inconspicuous in some genera (see table below)The face is elongated with coloring that sometimes has the appearance of the wasp mandibles. These flies have no mandibles. The abdomen is elongated often constricted in the first and/or second segments (see table below). The anterior half of the wing is commonly darkly colored while the posterior half is usually clear. The effect is the appearance of a dark forewing and a clear hind wing, as in wasps. When captured, the fly also has some behavioural mimicry, such as a stinging motion of the abdomen. These flies have no actual stinger. The overall coloration is very wasp-like, often black with yellow rings on the abdomen and strong markings on the thorax and face. Wing venation characteristics are of taxonomic value. The cross-vein r-m is at or beyond the middle of the discal cell. Cells r₁ and r₂₊₃ are open, while cell r₄₊₅ is closed near the margin of the wing. Vein R₄₊₅ can be straight, looping with or without a spur vein. ^[4] ^[5] ^[6]

Gallery

Wasp-mimic Ceriana ornata_
Ceriana alvesgaspar
Frontal prominence of Ceriana
Cerioidini antennae diagram
wing veination
Sphiximorpha subsessilis
Polybiomyia_bellardii

Genera characteristics

Genera of Cerioidini ^[6] ^[5]
Genus	frontal prominence	abdominal segment 1	abdominal segment two constriction	abdominal segment 2 length	paraface	other
Ceriana	as long as scape	same size as segment 2	not constricted	wider than long	very short paraface	looping in third vein R4+5 with spur branch projecting into cell r4+5
Monoceromyia	as long as scape	wider than segment 2	strongly constricted	longer than wide		loop of vein R4+5 without Spur branch
Polybiomyia	absent or very short	shorter than segment 2	strongly constricted	longer than wide		vein R4+5 not looping
Primocerioides	shorter than scape		not constricted	wider than long	long paraface and large anterior tentorial pit	eyes and face pilose, vein R4+5 straight
Sphiximorpha	shorter than scape	wider than 2	slightly constricted anteriomedially	longer than wide	long paraface and large or deep anterior tentorial pit.	Facial tubercle weak to nearly absent. vein R4+5 not looping.

References

^ Ricarte, A., Marcos-garcía, M. A., Rotheray, G. E. & Hancock, E. G. (2007). The Early Stages and Breeding Sites of 10 Cerioidini Flies (Diptera: Syrphidae). Annals of the Entomological Society of America 100(6):914-924 doi:10.1603/0013-8746(2007)100[914:TESABS2.0.CO;2]
^ Vockeroth, J.R. (1987). Manual of Nearctic Diptera, Volume 2 (PDF). Canada: Research Branch, Agriculture Canada. p. 668. ISBN 0-660-12125-5.
^ Heiss, Elizabeth Madeleine (1938). "A classification of the larvae and puparia of the Syrphidae of Illinois exclusive of aquatic forms". Series: Illinois Biological Monographs. 16: 1–142. end of template cite journal
^ Ferguson, E. W. (1926). "Revision of Australian Syrphidae (Diptera). Part i." (PDF). Proceedings of the Linnean Society of New South Wales. 51: 137–183.
^ ^a ^b Brunetti, Enrico (1923). Diptera Vol. III. Pipunculidae, Syrphidae, Conopidae, Oestridae. Vol. 3. London, England: Taylor and Francis. pp. 1–424.
^ ^a ^b Shannon, Raymond (1927). "Notes on and descriptions of Syrphid flies of the subfamily Cerioidinae". Journal of the Washington Academy of Sciences. 17: 38–53.

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[1] Ricarte, A., Marcos-garcía, M. A., Rotheray, G. E. & Hancock, E. G. (2007). The Early Stages and Breeding Sites of 10 Cerioidini Flies (Diptera: Syrphidae). Annals of the Entomological Society of America 100(6):914-924 doi:10.1603/0013-8746(2007)100[914:TESABS2.0.CO;2]

[2] Vockeroth, J.R. (1987). Manual of Nearctic Diptera, Volume 2 (PDF). Canada: Research Branch, Agriculture Canada. p. 668. ISBN 0-660-12125-5.

[Heiss-3] Heiss, Elizabeth Madeleine (1938). "A classification of the larvae and puparia of the Syrphidae of Illinois exclusive of aquatic forms". Series: Illinois Biological Monographs. 16: 1–142. end of template cite journal

[ferg26-4] Ferguson, E. W. (1926). "Revision of Australian Syrphidae (Diptera). Part i." (PDF). Proceedings of the Linnean Society of New South Wales. 51: 137–183.

[Brunetti-5] Brunetti, Enrico (1923). Diptera Vol. III. Pipunculidae, Syrphidae, Conopidae, Oestridae. Vol. 3. London, England: Taylor and Francis. pp. 1–424.

[shannon-6] Shannon, Raymond (1927). "Notes on and descriptions of Syrphid flies of the subfamily Cerioidinae". Journal of the Washington Academy of Sciences. 17: 38–53.

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